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the main thing to know here is "They concede they had known little about the nightclub scene". when you know little you open yourself up to get this kinda of treatment.
i imagine 90% of the tips he shelled out where just on the notion he "had" to. to break it down table reservations get treated the best as far as the average nightclub patron goes. no reason to tip the door guy when you are reserved, hes there to look for handouts...dont give it to him. tipping all the people on the way up was unnecessary and ive never seen anyone do it.
almost all nightclubs charge a 20 (not28)% gratuity and its included in the bill thats why they charged around 500 instead of 375. so giving the chick 50 bucks was also unnecessary.
the bathroom thing i can understand the lines get long. but ive never seen anyone tip ppl to go ahead of others. easy solution is to know that there are 2 bathrooms upstairs that are barely occupied, so tough luck there.
its all about knowing your environment. and if i could offer YOU a tip it would be dont tip anyone who asks for it, they probably dont deserve it.